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The gameplay features the return of Dante and Nero as playable characters, along with a new character, named "V". The gameplay is similar to the other titles in the Devil May Cry series, focusing on fast-paced "stylish action". The player fights off hordes of demons with a variety of attacks and weapons and receives a style-rating in combat based on a number of factors, such as move variety, the length of a combo and dodging attacks. The music in the game will change based on the player's performance in combat.
Nero is equipped with his Red Queen sword, his Blue Rose double-barreled revolver, and an assortment of new robotic arms called Devil Breakers, with a variety of functions such as grabbing enemies from a distance or stopping time to freeze an enemy in place. Along with his signature blade Rebellion and the demonic sword Sparda, Dante was shown using two new Devil Arms, a pair of buzzsaw-like weapons that combine into a motorcycle called Cavaliere, and a set of fiery gauntlets and boots called Balrog. In addition, Dante also wield Cerberus, introduced in Devil May Cry 3.The third playable character, V, uses three demons based on enemies from the first game to fight which includes Griffon, an eagle that uses ranged lightning based attacks, Shadow, a panther that forms blades, spikes and portals out of its body, and Nightmare, a large golem which V enters a Devil Trigger-like state, which turns his hair white, to summon, which uses a combination of melee attacks and laser beams.